A good stand and uniform plant population is a critical step in the crop production process. Crops that require expensive seeds and have tight production schedules require extra diligence in securing a good stand, which can be challenging. A review of
the fundamental stages of seed germination and seedling development can be helpful in early season field evaluations and management.
Seeds are living organisms, and they need healthy embryonic tissue to germinate. Most plant species have food reserves to carry the seedling through germination and emergence. The embryo and food reserves are covered in a seed coat and the seeds are stimulated,
or they “wake up”, in response to appropriate moisture, temperature, and light conditions that serve to initiate the germination process (Moore et al., 1998).
There are three distinct steps in the process of seed germination.
Several cool season plant species, including lettuce experience thermodormancy (Reynolds and Thompson, 1971), which is a state of dormancy induced by high temperatures that prevent germination. This is an adaptive trait that protects seeds from sprouting
during hot conditions that can create plant stress and poor-quality growth (Gardner et al., 1985 and Maynard and Hochmuth, 2007).
A range of optimum temperatures for a select group of common vegetable crops is shown in Table 1. Lettuce has an optimum temperature range for germination of 40-80°F with an optimum temperature of 75°F and a maximum temperature of 85°F.
The soil temperature conditions commonly experienced in the low deserts in August, September, and October when thousands of acres of lettuce are being planted and established, commonly exceed these lettuce temperature ranges for germination.
Table 1. Crop germination temperature ranges for a select set of vegetable crops. Source: Kemble and Musgrove, 2006.
The temperature limits for lettuce have been modified to some extent by plant breeding and improvement programs that have produced lettuce varieties that can better accommodate hot conditions. However, there are limits to a plant’s genetic elasticity in this regard. Development of some genetic tolerance to heat is being accomplished in crops such as lettuce with modern crop improvement programs. But we must realistically recognize that we are not changing a cool-season plant to a warm-season plant.
Several aspects of crop and soil management can help mitigate the extreme effects of heat on the soil surface. Perhaps the best and most common is the use of sprinkler irrigation systems to both moisten the soil and cool surface temperatures to a range that encourages lettuce seed germination and emergence (Figure 1).
Figure 1. Sprinkler irrigation on a recently planted lettuce field, Yuma Valley, Arizona.
The first step in germination is water entering the plant (imbibing) water from the soil. In dry lettuce seeds for example, this is primarily a physical process driven by the high-water potential gradient between the dry seed and the surrounding moist
soil medium. For many seeds, including lettuce, a key entry point for water is the micropyle, which is a small pore or opening in the seed coat. In many species, water enters through the micropylar end, which is where the radicle (embryonic root)
will later emerge.
The first thing to emerge from the seed is the primary root, called the radicle, which we often refer to as the “stinger” (Figure 2). The primary root serves to both anchor the plant to the ground and it also begins to absorb water.
After enough water is absorbed into the seed, the shoot emerges. In dicot plants (most broadleaf plants), the shoot has three main parts: the cotyledons (seed leaves), the hypocotyl, the section of shoot below the cotyledons, and the epicotyl, the section of shoot above the cotyledons (DuPont, 2025).
Figure 2. Germination stages for lettuce and other dicotyledonous crops. Source: Seeds and Seedling Biology, Penn. State University, 2025.
These early stages of germination and emergence shown in Figure 2 are critical in establishing a healthy plant, a strong stand, and a good plant population. Root development includes an elongation of the radical and the formation of lateral and secondary roots. The epicotyl will be elongating simultaneously, proceeding towards soil emergence (Figure 3).
Figure 3. Epigeal and hypogeal emergence. Source: Michael Knee, The Ohio State University. Further illustrations of emergent lettuce plants are shown in Figures 4 and 5.
During the emergence process when a dicot plant is in the “hook” stage, it is a critical time when the hook is trying to push through the soil surface. Plant species with small seeds are commonly not very strong with this process and that can be exacerbated with any soil crusting. This is another reason sprinkler irrigation is often used to diminish soil crusting and facilitate germination and emergence in the low desert vegetable crop production areas.
Figure 4. Lettuce cotyledons and first true leaves. Source: Harvest to Table.
Figure 5. Stages of emergence. Source: Spider Farmer.
Planting cool season crops, i.e., leafy green vegetables, in the lower Colorado River Valley in August, September, and October is a challenging operation. Seedling germination and early development are critical stages of growth that have lasting impact on the vigor and health of the crop.
References:
DuPont, T. Seed and Seedling Biology. 2025. Pennsylvania State University Cooperative Extension, ART 3208.
Gardner, Pearce, and Mitchell. 1985. Physiology of Crop Plants. The Iowa State University Press.
Kemble, J., and M. Musgrove. Soil Temperature Conditions for Vegetable Seed Germination. Alabama Cooperative Extension, 2006.
Knee, M. Development of Seed to Plant. 2024. The Ohio State University.
Maynard, D., and G. Hochmuth. Knott's Handbook for Vegetable Growers. Vol. 5. Hoboken, N.J.: John Wiley and Sons, 2007.
Moore R., W.D. Clark, and D.S. Vodopich. Botany. The McGraw-Hill Companies. 1998. ISBN: 0-697-38363-1
Reynolds, T. and P.A. Thompson.1971. Characterization of the high temperature inhibition of germination in lettuce (Lactuca sativa). Physiologia P1.24:544-547.
Fall melon season is approaching, and one recurring question I've been hearing is: Will viruses be as bad this fall as they were in the spring?
The incidence and severity of melon viruses this past spring were unprecedented across Yuma County, Imperial County, and northern Mexico. This is supported by the volume of feedback we received from growers, PCAs, and industry representatives who attended the June 2nd melon virus incident response meeting. As a result, predicting what we can expect is going to happen this fall is difficult. We have no recent, if any, experience with virus pressure at this scale in spring melon to guide our expectations for the upcoming fall season. At this point, predictions are more of an educated guess without the guidance of past observations.
To quickly recap, the three main viruses that affected cucurbits this spring were cucurbit yellow stunting disorder virus (CYSDV), cucurbit chlorotic yellows virus (CCYV), and watermelon chlorotic stunt virus (WmCSV). All three are transmitted by the Biotype B whitefly (Bemisia tabaci), whose populations overwintered at unusually high levels between 2025 and 2026. Between melon seasons, these viruses persist in a wide range of crop and weed hosts, many of which show few or no visible symptoms of infection. Unfortunately, these asymptomatic plants can still serve as reservoirs, allowing both the viruses and their whitefly vectors to bridge the melon-free gap between cropping seasons and provide a source of inoculum for newly planted fields. It is an unfortunate reality that neither the whitefly vectors nor many of the alternate host plants (weeds) show symptoms or suffer ill effects while carrying these viruses. As a result, they can stealthily maintain virus populations between melon seasons and serve as a source of infection for newly planted fields.
Below is a compilation of reported host plants for CYSDV, CCYV, and WmCSV. This list reflects the viruses’ confirmed hosts identified to date but is unlikely to be exhaustive. Additional weed and crop species may also be capable of serving as reservoirs for these viruses but have yet to be discovered or reported. Note that many of these plants may grow throughout the region as weeds, native vegetation, commercial crops, or in backyard gardens:
Table 1: Primary and alternate hosts of CYSDV, CCYV, and WmCSV reported to date.
I can see this upcoming melon season unfolding in one of two ways. On one hand, the most intuitive prediction is that the high virus inoculum and abundant whitefly populations present during the spring melon season will carry over into the fall, resulting in early and significant virus pressure. On the other hand, the intensive whitefly management programs implemented throughout the spring may have suppressed vector populations to provide knockdown to pre-winter 2025 levels, resulting in lower virus incidence early in the season than at the start of spring.
Regardless of which scenario plays out, proactive and preventative management of both whiteflies and weed reservoirs remains the most effective strategy for minimizing virus pressure in fall melons. This approach targets two critical stages of the disease cycle by reducing the initial sources of virus inoculum and limiting the whitefly vectors responsible for further virus spread.
Dr. Palumbo developed a management guide for whiteflies and CYSDV in fall melons in 2024. The recommendations are based on research findings from two key publications and provide practical guidance for reducing virus risk throughout the season, from planting through netted fruit (Castle 2017a and 2017b).
Table 2: Insecticide Use Guidelines for Whitefly /CYSDV Management in Fall Melons
The earlier melons become infected with one or more of these viruses, the greater the impact on plant growth, fruit development, and ultimately yield. Even when infection cannot be completely prevented, delaying virus establishment can substantially reduce losses in both yield and fruit quality. Protecting young plants from early whitefly feeding, and virus infection, is therefore one of the most important management objectives to reducing losses.
In field trials comparing at-plant soil applications of flupyradifurone (trade name Sivanto), dinotefuran (Venom), imidacloprid (Admire Pro), and cyantraniliprole (Verimark), Dr. Palumbo and colleagues found that flupyradifurone and dinotefuran provided the greatest protection against both whiteflies and lowest final incidence of CYSDV (Castle et al. 2017b). All products were applied as a single soil shank injection at planting, allowing systemic uptake and protection during crop establishment.
Across both spring and fall trials, flupyradifurone consistently produced the lowest whitefly densities and the lowest incidence of CYSDV. Dinotefuran was the second most effective treatment, significantly reducing both whitefly populations and CYSDV incidence, although its performance was somewhat less consistent than flupyradifurone. In contrast, at-planting treatment with imidacloprid and cyantraniliprole did not consistently reduce CYSDV incidence.
Further reading:
Castle, S., Palumbo, J., Merten, P., Cowden, C. and Prabhaker, N. (2017a), Effects of foliar and systemic insecticides on whitefly transmission and incidence of cucurbit yellow stunting disorder virus. Pest. Manag. Sci., 73: 1462-1472. https://doi.org/10.1002/ps.4478
Castle, S.J., Palumbo, J.P., Merten, P. (2017b), Field evaluation of cucurbit yellow stunting disorder virus transmission by Bemisia tabaci. Virus Res., 241:220-227. doi: https://doi.org/10.1016/j.virusres.2017.03.017
If you have any concerns regarding the health of your plants/crops please consider submitting samples to the Yuma Plant Health Clinic for diagnostic service or booking a field visit with me:
Christopher Detranaltes, Ph.D.
Cooperative Extension – Yuma County
Email: cdetranaltes@arizona.edu
Cell: 602-689-7328
6425 W 8th St Yuma, Arizona 85364 – Room 109
It is with mixed emotions that I write to inform you that this will be my last University of Arizona Vegetable IPM Update. The reason – I am retiring. Thank you for your support over the years. It’s been an honor and a privilege working with you and serving this ag community.
Figure 1. Automated Thinning & Weeding Technologies Field Day.
(Photo credits: Rosa Bevington)
As lettuce operations are transitioning to drip irrigation rather than overhead sprinklers to maximize water efficiency in the lower Colorado River, farm management practices must adapt, especially weed control. Pre-emergence tools like Prefar 4-E (bensulide) and Kerb SC (propyzamide) remain vital for managing purslane, chenopodium species, annual bluegrass, and key broadleaves. However, both active ingredients depend on specific soil placement and water delivery to form an effective barrier against germinating weed seeds.
Historically, standard overhead sprinkler passes provided the uniform surface coverage and water volume needed to activate Prefar and wash Kerb SC into the target seed zone. As fields transition to surface or subsurface drip irrigation, moving away from overhead water creates a distinct field challenge: drip lines push moisture laterally and downward, often leaving the top 1 to 2 inches of soil too dry to activate surface-applied herbicides.
Matching the Tool to the Mechanism: Because these two herbicides interact with soil and water differently, switching to drip requires tailored incorporation strategies:
Insect pests are constantly adapting to their environment. Over time, they have evolved ways to overcome control tactics. Repeated use of the same insecticides or tactics places strong selection pressure on pest populations. This allows the few individuals
that can survive treatments to reproduce, leading to populations that are increasingly difficult to control, a process known as resistance.
Insects can develop resistance in several ways. Some break down insecticides more efficiently or develop changes at the target site that reduce product effectiveness. Others avoid exposure altogether by changing their feeding habits or movement patterns. In some cases, insects even develop thicker cuticles that reduce or slow down the absorption of insecticides.
Production practices can influence the rate of resistance development. Large, uniform cropping systems and repeated use of the same control tools reduce opportunities for susceptible insects to persist, accelerating resistance.
Figure 1: Chart illustrating how insecticide resistance develops when insecticides with the same modeof action (MoA) are used repeatedly.
What Should You Do to Mitigate Resistance in Insect Pests?
An integrated pest management (IPM) approach remains the best defense against resistance.
Key practices include:
- Using crop varieties that are resistant to pest attack
- Rotating insecticides with different modes of action (MoA)
- Avoiding unnecessary insecticide applications
- Preserving beneficial insects by selecting insecticides with no or minimal adverse effects on them
- Incorporating non-chemical management tools whenever possible
- Maintaining refuges to sustain susceptible pest populations
- Eliminating crop residues after harvest to remove food sources and breeding sites for pests.
No insecticide remains fully effective over the long-term if overused. Reducing selection pressure by avoiding consecutive applications of insecticides from the same IRAC group is critical. The more intensively a single tactic is used, the faster resistance will develop. Resistance can increase production costs, reduce marketable yield, and ultimately affect grower competitiveness. A diversified pest management approach helps preserve the effectiveness of available tools and supports long-term, sustainable pest control.
Additional Reading Materials
- Calvin W., M. N. Keith, and B. McGrew. 2025. Guidelines for effective management of diamondback moth in brassica crops. University of Arizona Extension Publication. az2143. https://extension.arizona.edu/publication/guidelines-effective-management-diamondbackmoth-brassica-crops
- Calvin W. 2026. Effects of the Maximum Dose of Common Insecticides and Incipio on Diamondback Moth Populations Collected from Arizona and California Brassica Crops (Fall 2025– Winter/Spring 2026). https://acis.cals.arizona.edu/agriculturalipm/vegetables/vipm-archive/vipm-specialty-crops-view/effects-of-the-maximum-doseof-common-insecticides-and-incipio-on-diamondback-moth-populations-collected-fromarizona-and-california-brassica-crops-(fall-2025-winter-spring-2026)
Yuma is heading into fall produce season after an unusually warm year. By July 31, 2026, the season had accumulated 553 heat units (86/55°F) more than the 2020–2025 average. Through mid-August, nighttime temperatures were the warmest for that period in the AZMet Yuma Valley record, with 15 of 17 nights never dropping below 80°F. So the simple question is: if we go into produce season with above-average days and above-average nights, what should we expect? So, the useful answer starts with a distinction. Warm days and warm nights are not the same problem. They affect different parts of the crop, on different timelines, and they call for different management responses.
Warm Days, Part One: A Water-Demand Problem You Can Calculate Hot afternoons raise vapor pressure deficit and pull more water from the soil and crop. But the size of that effect is smaller than many people assume, and it is predictable. Averaged over 2020–2025, standardized reference evapotranspiration (ETo) during the establishment period was 1.69 inches per week in September, 1.26 inches in October, and 0.93 inches in November, a 45% decline across the window. Meanwhile, the spread between the warmest and coolest September in that record was only about one inch over the whole month. That comparison is the point: planting date moves water demand far more than how hot the year is. A warm September is not a reason to add a blanket percentage to every irrigation set. It is a reason to watch ETo and soil moisture closely during the two or three weeks when the crop has a limited root system and no canopy to protect the bed.
Warm Days, Part Two: A Germination Problem
This is the issue that can cost stands, and it is easy to miss because it looks like a water problem. Lettuce germination can be inhibited above roughly 81°F, and the effect is
stronger in darkness, which is exactly where the seed is. Prolonged exposure can push imbibed seed into secondary dormancy, at which point cooling the bed may no longer rescue it. Mean September soil temperature at 4 inches at the AZMet Yuma Valley
station over the past six years was 85, 85, 86, 87, 89, and 93°F. Every year was above the threshold. In a warm year, the margin gets worse rather than better. More water alone does not fix this. Variety selection, seed priming,
planting during the coolest part of the day, and sprinkler cooling of the seed zone are more targeted tools. Also, station soil temperature is a regional index, not the temperature in your bed. A bare, tilled, sunlit seedbed can run warmer. Put a
thermometer in the bed at seeding depth and decide on that number.
Warm Soils and Surface Drying
Warm conditions also affect the seedbed and root zone. Faster surface drying can concentrate salts near the seed line, especially where moisture is not uniform. This matters during lettuce establishment
because small seedlings have limited roots and are sensitive to both heat and salinity stress. The goal is not simply more water, but more uniform moisture in the right zone. Bed temperature, seed-zone moisture, and salt distribution should be checked
together before and after planting.
Warm Nights: A Marketability and Quality Problem
Nights get less attention than hot afternoons, but this year they are the more distinctive signal. Lettuce spends energy overnight on maintenance respiration. When nights stay warm,
more of the day’s photosynthate is burned rather than stored, and there is less recovery time after daytime stress. Later in the crop cycle, that can affect head development, firmness, uniformity, and tip burn risk. The cost lands on marketable
yield. Lettuce is bought on quality, and stress that reduces uniformity or head structure can cost more than the water it appeared to save. The encouraging part is that nighttime minimum temperatures usually fall from about 74°F in September to
about 62°F in October, so this pressure eases quickly with planting date.
Both Together: Pest Timing Can Move Up
Insect development is influenced by accumulated heat, so a season running 553°F-days ahead can also move pest timing ahead. That does not necessarily mean higher pest pressure; it means
the timing may shift.
What to Expect, in Short
Notice that four of those five pressures ease substantially between September and October. That is the practical thread running through all of this: in a warm year, planting date is one of the most powerful tools available, and the two weeks around establishment are where precision pays. A hot start does not call for guesswork, and it does not automatically call for more water or more inputs. Yuma growers have built their success by responding to heat with precision: better timing, closer measurement, stronger scouting, and careful root-zone management before produce season begins.
